List of Scarabaeidae subfamilies and tribes
The beetle family Scarabaeidae is made up of about 28 subfamilies containing more than 100 tribes. There are more than 2,500 genera and 35,000 described species in Scarabaeidae.[1]
The following subfamilies and tribes are in accordance with those in Catalog of Life (2023) and Dietz, Ahrens, et al. (2023). The tribe Hopliini is placed as a subfamily in Catalogue of Life, but is listed as a tribe of subfamily Melolonthinae here. The subfamilies Sericinae and Sericoidinae were suggested in Dietz, Ahrens, et al. (2023) and are included in the Catalogue of Life.[1][2] The subfamily Lichniinae is sometimes considered a tribe, Lichniini.[3]
Subfamily Aclopinae Blanchard, 1850
- Aclopini Blanchard, 1850
- Holcorobeini Nikolajev, 1992
- Phaenognathini Iablokoff-Khnozorian, 1977
Subfamily Aegialiinae Castelnau, 1840
- Aegialiini Castelnau, 1840
- Saprinianini Nikolajev, 2011
Subfamily Allidiostomatinae Arrow, 1940
Subfamily Aphodiinae Leach, 1815
- Aphodiini Leach, 1815
- Corythoderini Schmidt, 1910
- Eupariini Schmidt, 1910
- Hornietiini Minkina, 2020
- Odochilini Rakovič, 1987
- Odontolochini Stebnicka & Howden, 1996
- Psammodiini Mulsant, 1842
- Rhyparini Schmidt, 1910
- Stereomerini Howden & Storey, 1992
- Termitoderini Tangelder & Krikken, 1982
- Thinorycterini Semenov & Reichardt, 1925
- †Psammaegialiini Nikolajev, Wang & Zhang, 2014
Subfamily Aulonocneminae Janssens, 1946
Subfamily Cetoniinae Leach, 1815
- Cetoniini Leach, 1815
- Cremastocheilini Burmeister & Schaum, 1841
- Diplognathini Burmeister, 1842
- Goliathini Latreille, 1829
- Gymnetini Kirby, 1827
- Phaedimini Schoch, 1894
- Schizorhinini Burmeister, 1842
- Stenotarsiini Kraatz, 1880
- Taenioderini Mikšić, 1976
- Xiphoscelidini Krikken, 1984
Subfamily Chironinae Blanchard, 1845
Subfamily Dynamopodinae Arrow, 1911
Subfamily Dynastinae MacLeay, 1819
- Agaocephalini Burmeister, 1847
- Cyclocephalini Castelnau, 1840
- Dynastini Macleay, 1819
- Hexodontini Lacordaire, 1855
- Oryctini Mulsant, 1842
- Oryctoderini Endrödi, 1966
- Pentodontini Mulsant, 1842
- Phileurini Burmeister, 1847
Subfamily Eremazinae Stebnicka, 1977
- Eremazini Stebnicka, 1977
- †Yixianscarabaeini Nikolajev, 2015
Subfamily Lichniinae Burmeister, 1844
Subfamily Melolonthinae MacLeay, 1819
- Acomini Evans & Smith, 2020
- Ardellini Paulsen, 2021
- Athliini Smith & Evans, 2018
- Chasmatopterini Lacordaire, 1855
- Chnaunanthini Evans & Smith, 2020
- Colymbomorphini Blanchard, 1850
- Comophini Britton, 1978
- Dichelonychini Kirby, 1837
- Diphycerini Medvedev, 1952
- Diplotaxini Kirby, 1837
- Euchirini Hope, 1840
- Hopliini Latreille, 1829
- Langbianellini Prokofiev, 2015
- Macrodactylini Kirby, 1837
- Melolonthini Leach, 1819
- Pachypodini Erichson, 1840
- Pachytrichini Burmeister, 1855
- Phobetusini Evans & Smith, 2020
- Phyllotocidiini Britton, 1957
- Rhizotrogini Burmeister, 1855
- Systellopini Dalla Torre, 1912
- Warwickiini Evans & Smith, 2020
- †Cretomelolonthini Nikolajev, 1998
Subfamily Oncerinae LeConte, 1861
Subfamily Orphninae Erichson, 1847
- Aegidiini Paulian, 1984
- Orphnini Erichson, 1847
Subfamily Pachydeminae Reitter, 1902
- Aegosthetini Lacroix, 2007
- Pachydemini Burmeister, 1855
Subfamily Phaenomeridinae Erichson, 1847
Subfamily Podolasiinae Howden, 1997
Subfamily Rutelinae MacLeay, 1819
- Adoretini Burmeister, 1844
- Alvarengiini Frey, 1975
- Anatistini Lacordaire, 1855
- Anomalini Streubel, 1839
- Anoplognathini MacLeay, 1819
- Geniatini Burmeister, 1844
- Rutelini MacLeay, 1819
Subfamily Scarabaeinae Latreille, 1802
- Ateuchini Perty, 1830
- Byrrhidiini Davis, Deschodt & Scholtz, 2019
- Coprini Leach, 1815
- Deltochilini Lacordaire, 1855
- Dichotomiini Pereira, 1954
- Endroedyolini Davis, Deschodt & Scholtz, 2019
- Epactoidini Rossini et al, 2022
- Epilissini Lansberge, 1874
- Epirinini Lansberge, 1874
- Eucraniini Burmeister, 1873
- Gymnopleurini Streubel, 1846
- Odontolomini Davis, Deschodt & Scholtz, 2019
- Oniticellini Kolbe, 1905
- Onitini Castelnau, 1840
- Onthophagini Streubel, 1846
- Parachoriini Tarasov, 2017
- Phanaeini Hope, 1838
- Scarabaeini Latreille, 1802
- Sisyphini Mulsant, 1842
Subfamily Sericinae Kirby, 1837
- Ablaberini Blanchard, 1850
- Diphucephalini Castelnau, 1840
- Sericini Kirby, 1837
Subfamily Sericoidinae Erichson, 1847
- Automoliini Britton, 1978
- Heteronychini Lacordaire, 1855
- Liparetrini Burmeister, 1855
- Maechidiini Burmeister, 1855
- Phyllotocini Burmeister, 1855
- Scitalini Britton, 1957
- Sericoidini Erichson, 1847
Subfamily Termitotroginae Wasmann, 1918
Subfamily Trichiinae Kolbe, 1897
- Trichiini Fleming, 1821
Subfamily Valginae Schenkling, 1922
- Valgini Mulsant, 1842
Subfamily †Cretoscarabaeinae Nikolajev, 1995
Subfamily †Lithoscarabaeinae Nikolajev, 1992
Subfamily †Prototroginae Nikolajev, 2000
- †Prototrox Nikolajev, 2000
References
- ^ a b "Scarabaeidae Latreille, 1802". Catalogue of Life. Retrieved 2023-09-22.
- ^ Dietz, Lars; Seidel, Matthias; Eberle, Jonas; Misof, Bernhard; et al. (2023). "A transcriptome-based phylogeny of Scarabaeoidea confirms the sister group relationship of dung beetles and phytophagous pleurostict scarabs (Coleoptera)". Systematic Entomology. 48 (4). doi:10.1111/syen.12602.
- ^ Mondaca, José (2023). "A checklist of the Scarabaeoidea (Coleoptera) of Chile with exemplar live-photographed". Zootaxa. 5285 (2). doi:10.11646/ZOOTAXA.5285.2.1.