Lebanese amber

Lebanese amber is fossilized resin found in Lebanon and its surroundings. It dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. It formed on what was then the northern coast of Gondwana, believed to be a tropical or subtropical zone in a temperate or hot climate.[1] It is the oldest source of amber with a significant number of inclusions. Up to 300 sources of Lebanese amber have been recovered and 17 of them are important sources of organic inclusions, which are the oldest of their kind. The inclusions help to document Cretaceous fauna and flora.

Origins

Lebanese amber can be found in Lebanon and neighboring areas of the Levant. Up to 300 different sources of amber had been discovered by 2010. The amber was deposited in the Cretaceous era and is rich in fossil synclusions. 19 of the discovered sources are rich in inclusions from the Early Cretaceous. All of them are located in Lebanon, which makes it the largest source of inclusion from that period.[2]

History

Aside from possible early reports of Phoenician usage, the oldest reports of Lebanese amber are from 19th-century accounts, these tended only to be incidental due to Lebanese amber's gemological quality in comparison to Baltic amber, and the local people were more interested in the associated lignite as a source of fuel.[3]

Properties

Lebanese amber can be found in a vast variety of colors such as yellow, orange, dark red or iridescent jet black. Rarely in white, milky or cream. The variation of color tone is caused by the air contained in the amber. The density of Lebanese amber is 1.054 g/cm3. It tends to be fragile and easy to damage.[4]

Inclusions

Inclusions are quite common for Lebanese amber, which is rich in organisms of the Barremian epoch. Next to Jordanian amber, Lebanese amber is the oldest amber to have yielded significant invertebrate inclusions alongside the Wealden amber from the equivalently aged Wessex Formation of the UK, which is much less productive. Organisms preserved in Lebanese amber are dated back to the period prior to the angiosperm radiation, which was the period of massive extinction of old groups of arthropods, as well as the emergence of the new ones, some of which co-evolved with angiosperms. The organisms are preserved in good condition and shape.[3] The diversity and number of co-inclusions help to draw conclusions about mutual relations and co-existence.[5]

Paleobiota

After Poinar & Milki (2001),[1] Maksoud & Azar (2020)[6] and subsequent studies.

Bacteria

Taxon Authority Year described Notes Images

Streptomyces sp.

Fungi

Taxon Authority Year described Notes Images

Chytridiales indet.

Mucorales indet.

Plants

Taxon Authority Year described Notes Images

Agathis levantensis

Poinar & Milki 2001 An araucarian tree responsible for the production of amber.

Hexapoda

Taxon Authority Year described Notes Images

Archaeatropos

Azar & Nel 2004 An archaeatropid psocodean.[7]

Arthropleona indet.

A springtail.

Asphaeropsocites

Azar et al. 2010 A sphaeropsocid psocodean.[8]

Aphelopus

Olmi 1998 A dryinid wasp.

Archiaustroconops

Szadziewski 1996 A ceratopogonid fly.

Archiculicoides

Szadziewski 1996 A ceratopogonid fly.

Archisciada

Grimaldi & Cumming 1999 A sciadocerid fly.

Atelestites

Grimaldi & Cumming 1999 An empidid fly.

Austroconops

Szadziewski 1996 A ceratopogonid fly.

Avenaphora

Grimaldi & Cumming 1999 A dolichopodid fly.

Banoberotha

Whalley 1980 A beaded lacewing.

Bcharreglaris

Azar & Nel 2004 An archaeatropid psocodean.[7]

Bernaea

Schlee 1970 A whitefly.

Blattodea indet.

A cockroach.[8]

Chomeromyia

Grimaldi & Cumming 1999 A fly.

Conovirilus

McCafferty 1997 A leptophlebiid mayfly.

Corethrella

Szadziewski 1995 A corethrellid biting midge.

Cretaceomachilis

Sturm & Poinar, 1998 A meinertellid bristletail.

Cretacetrocta

Hakim & Azar 2024 A pachytroctid psocodean.[7]

Cretapsychoda

Azar et al. 1999A A psychodid fly.

Enicocephalinus

Azar et al. 1999B An enicocephalid.

Exitelothrips

zur Strassen 1973 A scudderothripid thrips.

Fossileptoconops

Szadziewski 1996 A ceratopogonid fly.

Glaesoconis

Whalley 1980 A dustywing.

Heidea

Schlee 1970 A whitefly.

Incurvariites

Whalley 1978 A incurvariid moth.
Jankotejacoccus Szwedo, Azar & Sendi 2025 A jankotejacoccid scale insect.[9]

Jezzinothrips

zur Strassen 1973 A jezzinothripid thrips.

Lebambromyia

Grimaldi & Cumming 1999 A phorid fly.

Lebanaphis

Heie & Azar 2000 A tajmyraphidid hemipteran.

Lebania

Podenas et al. 2001 A tipulid crane fly.

Lebanoconops

Szadziewski 1996 A ceratopogonid fly.

Lebanoculicoides

Szadziewski 1996 A ceratopogonid fly.

Leptoconops

Borkent 2000 A ceratopogonid fly.

Libaneuphoris

Azar, Huang, Cai & Nel 2015 A libanophorid pscocodean.

Libanobythus

Prentice et al. 1996 A scolobythid hemipteran.

Libanochlites

Brundin 1976 A non-biting midge.

Libanoculex

Azar, Nel, Huang & Engel 2023 A chaoborid fly, previously interpreted as a mosquito.[10]

Libanoglaris

Azar & Nel 2004 An archaeatropid psocodean.[7]

Libanophlebotomus

Azar et al. 1999A A psychodid fly.

Libanomphientomum

Choufani, Azar & Nel 2011 A electrentomoid psocodean.[7]

Libanopsychoda

Azar et al. 1999A A psychodid fly.

Libanopsyllipsocus

Azar & Nel 2011 A psyllipsocid psocodean.[7]

Libanorhinus

Kuschel & Poinar 1993 A nemonychid weevil.

Libanosemidalis

Azar et al. 2000 A dustywing.

Lonchopterites

Grimaldi & Cumming 1999 A lonchopterid fly.

Lonchopteromorpha

Grimaldi & Cumming 1999 A lonchopterid fly.

Rhadinolabis

Engel, Ortega-Blanco & Azar 2011 An earwig.[8]

Setoglaris

Azar & Nel 2004 An archaeatropid psocodean.[7]

Lebanotermes

Engel, Azar & Nel in Engel et al. 2011 A termite.[8]

Megarostrum

Heie & Azar 2000 A tajmyraphidid hemipteran.

Mesobolbomyia

Grimaldi & Cumming 1999 A rhagionid fly.

Mundopoides

Fennah 1987 A cixiid.

Neocomothrips

zur Strassen 1973 A neocomothripid thrips.

Paleochrysopilus

Grimaldi & Cumming 1999 A rhagionid fly.

Paleopsychoda

Azar et al. 1999A A psychodid fly.

Palaeosiamoglaris

Azar, Huang & Nel 2017 A prionoglarid psocodean.[7]

Parasabatinca

Whalley 1978 A micropterigid moth.

Paraberotha

Whalley 1980 A beaded lacewing.

Paramesopsocus

Azar, Hajar, Indary & Nel 2008 A electrentomid psocodean.[7]

Phaetempis

Grimaldi & Cumming 1999 An empidid fly.

Phlebotomites

Hennig 1972 A phlebotomid fly.

Progonothrips

zur Strassen 1973 A rhetinothripid thrips.

Protoculicoides

Szadziewski 1996 A ceratopogonid fly.

Protopsychoda

Azar et al. 1999A A psychodid fly.

Psocodea sp. 1

A juvenile psocodean.[7]

Raptorapax

Petrolevičius, Azar & Nel 2010 A rhachiberotid neuropteran.[8]

Rhetinothrips

zur Strassen 1973 A rhetinothripid thrip.

Scaphothrips

zur Strassen 1973 A scaphothripid thrips.

Scudderothrips

zur Strassen 1973 A scudderothripid thrips.

Sphaeropsocites

Grimaldi & Engel 2006 A sphaeropsocid psocodean.[7]

Trichinites

Hennig 1970 An empidid fly.

Yuripopovina

Azar et al. 2011 A yuripopovinid hemipteran.[8]

Arachnida

Taxon Authority Year described Notes Images

Acari indet.

Various free-living mites.

Anystidae sp.

An anystid mite.

Archaeobuthus

Lourenço 2001 An archaeobuthid scorpion.[8]

Erythraeoidea indet.

A erythraeid mite.

Erythraeoidea indet.

Leptus sp.?

Lebansegestria

Wunderlich 2008 A segestriid spider.

Lebanoecobius

Wunderlich 2004 A oecobiid spider.

Linyphiidae sp.

A linyphiid spider.

Microsegestria

Wunderlich & Milki 2004 A segestriid spider.

Neoliodes

Arillo, Subías, Chaves Da Rocha & Azar 2019 A neoliodid mite.

Oonopidae sp.

An oonopid spider.

Palaeomicromenneus

(Penney) 2003 A deinopid spider.

Plumorsolus

Wunderlich 2008 A plumorsolid spider.

Pseudoscorpiones indet.

A pseudoscorpion.

Zamilia

Wunderlich 2008 A oecobiid spider.

Other invertebrates

Taxon Authority Year described Notes Images

Chilopoda indet.

Cretaciomermis

Poinar 2001 A nematode.

Electroxenus

Nguyen Duy-Jacquemin & Azar 2004 A polyxenid millipede.

Heleidomermis

Poinar, Acra & Acra 1994 A mermithid nematode.

Libanoxenus

Nguyen Duy-Jacquemin & Azar 2004 A polyxenid millipede.

Pupilloidea indet.

A possible pupillid snail.

Vertebrates

Taxon Authority Year described Notes Images

Aves indet.

A single feather and the only known bird remains of the biota.

Baabdasaurus

Arnolds et al. 2002 A reptile.

References

  1. ^ a b Raif Milki, George Poinar, Lebanese Amber: The Oldest Insect Ecosystem in Fossilized Resin, 2001. ISBN 978-0-87071-533-4
  2. ^ David I. Green, David Penney, Fossils in Amber: Remarkable Snapshots of Prehistoric Forest Life, Siri Scientific Press, 2011
  3. ^ a b Penney, David (2010). "Chapter 14: Lebanese Amber". Biodiversity of fossils in amber from the major world deposits. Siri Scientific Press. ISBN 978-0-9558636-4-6. OCLC 904772774.
  4. ^ Andrew Ross, Amber, Harvard University Press, 1998
  5. ^ George Poinar, Life in amber, Stanford University Press, 1992
  6. ^ Maksoud, Sibelle; Azar, Dany (2020-04-30). "Lebanese amber: latest updates". Palaeoentomology. 3 (2): 125–155. Bibcode:2020Plegy...3..125M. doi:10.11646/palaeoentomology.3.2.2. ISSN 2624-2834.
  7. ^ a b c d e f g h i j k Marina, Hakim; Dany, Azar (2024). "Contributions to the Palaeobiodiversity of Psocodea ('Psocoptera') from Lebanese Amber: A Review". Fossil Studies. 2 (3). doi:10.3390/fo (inactive 1 July 2025). ISSN 2813-6284. Archived from the original on 2024-12-03.{{cite journal}}: CS1 maint: DOI inactive as of July 2025 (link)
  8. ^ a b c d e f g Azar, D. (2012). Lebanese amber: a “Guinness Book of Records”. Annales Universitatis Paedagogicae Cracoviensis. Studia ad Didacticam Biologiae Pertinentia, 2(I), 44-60.
  9. ^ Peter, Vršanský; Hemen, Sendi; Júlia, Kotulová; Jacek, Szwedo; Martina, Havelcová; Helena, Palková; Lucia, Vršanská; Jakub, Sakala; L'ubica, Puškelová; Marián, Golej; Adrian, Biroň; Daniel, Peyrot; Donald, Quicke; Didier, Néraudeau; Pavel, Uher (2025-02-07). "Jurassic Park approached: a coccid from Kimmeridgian cheirolepidiacean Aintourine Lebanese amber". National Science Review. 12 (3). doi:10.1093/n. ISSN 2095-5138. Archived from the original on 2025-05-05.
  10. ^ Harbach, Ralph E. (2024-03-12). "Libanoculex intermedius is not a mosquito (Diptera: Culicidae): It is a chaoborid (Chaoboridae)". Zootaxa. 5424 (1): 139–144. doi:10.11646/zootaxa.5424.1.9. ISSN 1175-5334. PMID 38480294. Archived from the original on 9 August 2024. Retrieved 9 August 2024.